NERC GW4+ DTP PhD studentship for September 2027 Entry - Costly signalling and mate search in wild fiddler crabs Ref: 6048

About the project

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Supervisors

Lead Supervisor

Dr Tim Fawcett , Exeter, Psychology

Additional Supervisors

Dr Safi Darden, Exeter, Psychology

Dr Martin How, Bristol, Biological Sciences

Location: University of Exeter

About the Partnership

This project is one of a number that are in competition for funding from the NERC Great Western Four+ Doctoral Training Partnership (GW4+ DTP). The GW4+ DTP consists of the Great Western Four alliance of the University of Bath, University of Bristol, Cardiff University and the University of Exeter plus five Research Organisation partners: British Antarctic Survey, British Geological Survey, Centre for Ecology and Hydrology, the Natural History Museum and Plymouth Marine Laboratory. The partnership aims to provide a broad training in earth and environmental sciences, designed to train tomorrow’s leaders in earth and environmental science. For further details about the programme please see http://nercgw4plus.ac.uk/

For eligible successful applicants, the studentships comprises:

Project Aims and Methods

Sexual selection is a potent evolutionary force, responsible for astonishing diversity in morphology and behaviour. To understand how such striking displays have evolved, we need to know (1) how individuals adjust their display behaviour depending on the social environment and their own physiological state; and (2) how individuals choose among potential mates whose displays change rapidly over time. This PhD project will tackle these questions using a combination of evolutionary modelling and cutting-edge behavioural experiments in a wild population of West African fiddler crabs (Afruca tangeri). The theoretical component will involve building models (using dynamic programming, or evolutionary computer simulations, or both) to predict adaptive strategies for courtship and mate search. The empirical component will involve field trips (May-July) to our study site in southern Portugal, where thousands of fiddler crabs can be studied at close quarters in their natural environment. In this species, male crabs develop an enlarged major claw (around 30% of their body weight) that they use in waving and drumming displays to attract females. We have recently developed techniques to measure the energetic costs of male courtship in real time using flow-through respirometry, and to simulate competition from rival males using biomimetic robots. We can also follow the detailed mate-search behaviour of female crabs using miniature Bluetooth devices and automated video tracking. Our supervisory team is an interdisciplinary collaboration across the DLTP partnership and will provide bespoke training in behavioural fieldwork (Darden—Exeter), ecophysiology and sensory ecology (How—Bristol), game theory and evolutionary simulations (Fawcett—Exeter), as well as more general skills including data analysis and scientific writing. There are many exciting directions the work could take and we are especially keen to hear from students who want to play an active role in designing the project.

Further reading:

Wilde et al. (2025) https://doi.org/10.1098/rspb.2025.1570

Hollon et al. (2023) https://doi.org/10.1007/s00265-023-03296-9

Useful recruitment links:

Webpage : https://www.exeter.ac.uk/research/groups/psychology/crab/

For information relating to the research project please contact the lead Supervisor via: t.w.fawcett@exeter.ac.uk

• You will receive a stipend, which will start at £21,805 p.a. for the academic year 2026/27, and expected to increase annually to reflect inflation.
• Your university tuition fees will be paid by us (£5,238 for 2026/27, expected to increase annually to reflect inflation).
• A Research Training Support Grant of £11,000 (plus contributions from a project partner on some projects) which is used for project costs such as lab and field work and attendance at an international conference.
• A further budget of £1,000 (which can be increased to £2,000 on application) to attend specialist training courses.

Eligibility

NERC GW4+ DTP studentships are open to UK and Irish nationals who, if successful in their applications, will receive a full studentship including payment of university tuition fees at the home fees rate.

A limited number of full studentships are also available to international students which are defined as EU (excluding Irish nationals), EEA, Swiss and all other non-UK nationals. For further details please see the NERC GW4+ website.

Those not meeting the nationality and residency requirements to be treated as a ‘home’ student may apply for a limited number of full studentships for international students. Although international students are usually charged a higher tuition fee rate than ‘home’ students, those international students offered a NERC GW4+ Doctoral Training Partnership full studentship starting in 2027 will only be charged the ‘home’ tuition fee rate (which will be covered by the studentship). International applicants need to be aware that you will have to cover the cost of your student visa, healthcare surcharge and other costs of moving to the UK to do a PhD. More information on this is available from the universities you are applying to (contact details are provided in the project description that you are interested in.

The conditions for eligibility of home fees status are complex and you will need to seek advice if you have moved to or from the UK (or Republic of Ireland) within the past 3 years or have applied for settled status under the EU Settlement Scheme.

Equality, Diversity and Inclusion

The University of Exeter is committed to promoting and supporting equality, diversity, and inclusion within our working environments and is at the heart of all our activities. With over 27,000 students and 6,400 staff from 180 different countries we offer a diverse and engaging environment where our diversity is celebrated and valued as a major strength.

We actively encourage applicants with varied experiences and backgrounds and from all sections of the community regardless of age, race, ethnicity, sexual orientation, gender, religion, or disability status. We are committed to creating an inclusive culture where all members of our community are supported to thrive.

Whilst all applicants will be judged on merit alone, we particularly welcome applications from groups currently underrepresented within our postgraduate research student community. Reasonable adjustments are available for interviews and workspaces.

Entry requirements

Applicants should have obtained, or be about to obtain, a First or Upper Second Class UK Honours degree, or the equivalent qualifications gained outside the UK. Applicants with a Lower Second Class degree will be considered if they also have Master’s degree. Applicants with a minimum of Upper Second Class degree and significant relevant non-academic experience are encouraged to apply.

All applicants would need to meet our English language requirements by the start of the project http://www.exeter.ac.uk/postgraduate/apply/english/.

How to apply

Apply now

In the application process you will be asked to upload several documents. Please note our preferred format is PDF, each file named with your surname and the name of the document, eg. “Smith – CV.pdf”, “Smith – Cover Letter.pdf”, “Smith – Transcript.pdf”.

Note: Applicants can only apply for one PhD project across the GW4+ DLTP Partnership. Any additional applications received will be deemed unsuccessful.

You are also required to complete a DLTP Survey Applicants for PhDs 2027 for EDI data collection, which can be completed via the following link : DLTP Survey Applicants for PhDs 2027 - Online Surveys | Online surveys . This information will not be seen by the supervisory team.

You may also opt to complete an additional statement, although this is not requirement of the application process. The additional statement can be completed via the following link https://app.onlinesurveys.jisc.ac.uk/s/nercdtp/gw4-dltp-optional-additional-statement-2027

Reference information You will be asked to submit two references as part of the application process. If you are not able to upload your reference documents with your application please ensure you provide details of your referees. If you provide contact details of referees only, we will not expect receipt of references until after the shortlisting stage. Your referees should not be from the prospective supervisory team.

If you are shortlisted for interview, please ensure that your two academic referees email their references to the pgrapplicants@exeter.ac.uk , 7 days prior to the interview dates. Please note that we will not be contacting referees to request references, you must arrange for them to be submitted to us by the deadline.

References should be submitted by your referees to us directly in the form of a letter. Referees must email their references to us from their institutional email accounts. We cannot accept references from personal/private email accounts, unless it is a scanned document on institutional headed paper and signed by the referee.

All application documents must be submitted in English. Certified translated copies of academic qualifications must also be provided.

The closing date for applications is Thursday 7 January 2027 at 2359 GMT . Shortlisting will be completed between 9th January - 14th February 2027. Applicants invited to interview will be notified w/c 15 February 2027, with interviews scheduled for 1 to 5 March 2027.

For more information about the NERC GW4+ DPT please visit https://gw4plusdltp.ac.uk/

If you have any general enquiries about the application process please email pgrapplicants@exeter.ac.uk. Project-specific queries should be directed to the lead supervisor.

Data Sharing During the application process, the University may need to make certain disclosures of your personal data to third parties to be able to administer your application, carry out interviews and select candidates. These are not limited to, but may include disclosures to:

Such disclosures will always be kept to the minimum amount of personal data required for the specific purpose. Your sensitive personal data (relating to disability and race/ethnicity) will not be disclosed without your explicit consent.

• CV
• DLTP Personal Statement (Please use the template in the link provided). If the personal statement is not included your application will not be considered.
• Transcript(s) giving full details of subjects studied and grades/marks obtained. This should be an interim transcript if you are still studying.
• If you are not a national of a majority English-speaking country you will need to submit evidence of your current proficiency in English, please see the entry requirements for details.
• Two references
• the selection panel and/or management board or equivalent of the relevant programme, which is likely to include staff from one or more other HEIs;
• administrative staff at one or more other HEIs participating in the relevant programme.

Summary

Application deadline: | 7th January 2027

Value: | For eligible students the studentship will cover home tuition fees plus an annual tax-free stipend of at least £21,805 p.a for 3.5 years full-time.

Duration of award: | per year

Contact: PGR Admissions | pgrapplicants@exeter.ac.uk

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